My main question is why you can't just run the new GPUs on your current motherboard, RAM, and CPU. If you're currently on stock cooling you could pick up a Hyper 212 Plus, OC your processor (and a fair bit too) and run the GPUs on your board's dual PCI-E x16 slots.
Anyway, if you feel compelled to upgrade your platform in addition to your GPUs, and you've got $500 to do it, I'd say go for an i5 750, a P7P55D, and 4GB of DDR3 memory. That should come out to about $500, depending on the deals you find.
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"If dancing is sure to result in failure then you must dance anyway."
Lo Mein said that, and I think he knows a little more about dancing than I do pal, because he invented it! Then he perfected it so that no man could best him in the club of honor.
Then he used his prize money to buy two of every whore on Earth. Then he herded them all onto a boat, then they all beat the crap out of every single one of him.
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